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add depensation in operating model pellatomlinson_rla

Matthieu Authier requested to merge depensation into master

Added new arg to pellatomlinson_rla: Allee which is a list of two arg. One arg. is a probability of a female meeting a male, and the other is a level of male depletion corresponding to the probability. Depensation is modelled with a negative exponential functional form.

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